In 2371, the Enterprise is pulled into an elaborate scheme hatched by Dr. Tolian Soran, who is attempting to re-enter the spatial anomaly known as the Nexus. The chase ends at Veridian III and Picard beams down to confront him. Soran's plan requires the destruction of the Veridian star, which will alter gravity and steer the Nexus ribbon toward him. In one timeline, Soran succeeds in destroying the star, and both he and Picard are swept up by the Nexus.

Within the Nexus, Picard encounters an "echo" of Guinan, once herself a Nexus denizen. She leads him to Captain James T. Kirk, who enters the realm during the Enterprise-B's maiden voyage. Picard convinces Kirk to leave the Nexus to help him stop Soran. In an apparent temporal paradox, Kirk and Picard appear on Veridian III before Soran can destroy the star. The two starship captains prevent Soran's star-destroying missile from launching, causing it to explode on the pad and to kill the nearby Soran; the Nexus ribbon passes harmlessly overhead. However, Kirk is killed during the fighting.

While Picard confronts Soran, the Enterprise confronts the Duras sisters and is engaged by their ship. Although the Enterprise destroys Lursa's and B'Etor's ship, the Enterprise's warp coolant system is seriously damaged, building up to a warp core breach. Commander William Riker orders the crew evacuated to the saucer section and commences a saucer separation. The explosion that destroys the stardrive section damages the saucer, causing it to crash virtually uncontrolled onto Veridian III. The saucer is not salvageable.

After the Enterprise's surviving crew are rescued, Picard is given command of the Enterprise-E, a new Sovereign class starship.

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